A regulatory circuit controlling the dynamics of NFkappaB cRel transitions B cells from proliferation to plasma cell differentiation
Journal Title
Immunity
Publication Type
Journal Article
Abstract
Humoral immunity depends on efficient activation of B cells and their subsequent differentiation into antibody-secreting cells (ASCs). The transcription factor NFkappaB cRel is critical for B cell proliferation, but incorporating its known regulatory interactions into a mathematical model of the ASC differentiation circuit prevented ASC generation in simulations. Indeed, experimental ectopic cRel expression blocked ASC differentiation by inhibiting the transcription factor Blimp1, and in wild-type (WT) cells cRel was dynamically repressed during ASC differentiation by Blimp1 binding the Rel locus. Including this bi-stable circuit of mutual cRel-Blimp1 antagonism into a multi-scale model revealed that dynamic repression of cRel controls the switch from B cell proliferation to ASC generation phases and hence the respective cell population dynamics. Our studies provide a mechanistic explanation of how dysregulation of this bi-stable circuit might result in pathologic B cell population phenotypes and thus offer new avenues for diagnostic stratification and treatment.
Publisher
Cell Press
Research Division(s)
Immunology
PubMed ID
30850343
NHMRC Grants
NHMRC/1054925NHMRC/1058238
Terms of Use/Rights Notice
Refer to copyright notice on published article.


Creation Date: 2019-03-13 11:54:00
Last Modified: 2019-03-13 12:05:31
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