Positive Feedback Between PU.1 and the Cell Cycle Controls Myeloid Differentiation
Details
Publication Year 2013-08-09,Volume 341,Issue #6146,Page 670-673
Journal Title
SCIENCE
Publication Type
Journal Article
Abstract
Regulatory gene circuits with positive-feedback loops control stem cell differentiation, but several mechanisms can contribute to positive feedback. Here, we dissect feedback mechanisms through which the transcription factor PU.1 controls lymphoid and myeloid differentiation. Quantitative live-cell imaging revealed that developing B cells decrease PU.1 levels by reducing PU.1 transcription, whereas developing macrophages increase PU.1 levels by lengthening their cell cycles, which causes stable PU.1 accumulation. Exogenous PU.1 expression in progenitors increases endogenous PU.1 levels by inducing cell cycle lengthening, implying positive feedback between a regulatory factor and the cell cycle. Mathematical modeling showed that this cell cycle-coupled feedback architecture effectively stabilizes a slow-dividing differentiated state. These results show that cell cycle duration functions as an integral part of a positive autoregulatory circuit to control cell fate.
Publisher
AMER ASSOC ADVANCEMENT SCIENCE
Keywords
TRANSCRIPTION FACTOR PU.1; HEMATOPOIETIC STEM-CELLS; GENE-EXPRESSION; LEUKEMIA; PROGENITORS; FATES; MICE; PROLIFERATION; ACTIVATION; MACROPHAGE
Research Division(s)
Molecular Immunology
Terms of Use/Rights Notice
Copyright © 2013 American Association for the Advancement of Science. All Rights Reserved


Creation Date: 2013-08-09 12:00:00
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