Unconventional binding of Calmodulin to CHK2 kinase inhibits catalytic activity
Journal Title
Biochemical Journal
Publication Type
Sep 23
Abstract
Calmodulin (CaM) serves an essential role in eukaryotic cells as a Ca2+ sensor. Ca2+ binding leads to conformation changes in CaM that enable engagement of a repertoire of enzymes and the regulation of their catalytic activities. Classically, Ca2+-CaM binds to an inhibitory pseudosubstrate sequence C-terminal to the kinase domain in members of the Ca2+-CaM dependent protein kinase (CAMK) family, and relieves inhibition to promote catalytic activity. Here, we report an unexpected mechanism by which CaM can bind CHK2 kinase to inhibit its kinase activity. Using biochemical, biophysical, and structural mass spectrometry, we identify a direct interaction of Ca2+-CaM with the CHK2 kinase domain that suppresses CHK2 catalytic activity in vitro and identify K373 in CHK2 as crucial for cell proliferation in human cells following DNA damage. Our findings add direct suppression of kinase activity to the repertoire of CaM's functions, complementing the paradigmatic mechanism of promoting kinase activity through autoinhibitory domain sequestration.
Publisher
Portland Press
Keywords
Allosteric regulation; Calcium signalling; Calmodulin; Cell cycle; Kinases
Research Division(s)
Inflammation; Advanced Technology and Biology; Structural Biology
PubMed ID
41003242
Open Access at Publisher's Site
https://doi.org/10.1042/BCJ20253431
Terms of Use/Rights Notice
Refer to copyright notice on published article.


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