BH3-only Bcl-2 family member Bim is required for apoptosis of autoreactive thymocytes
Details
Publication Year 2002-02-21,Volume 415,Issue #6874,Page 922-926
Journal Title
NATURE
Publication Type
Journal Article
Abstract
During lymphocyte development, the assembly of genes coding for antigen receptors occurs by the combinatorial linking of gene segments. The stochastic nature of this process gives rise to lymphocytes that can recognize self-antigens, thereby having the potential to induce autoimmune disease. Such autoreactive lymphocytes can be silenced by developmental arrest or unresponsiveness (anergy)(1), or can be deleted from the repertoire by cell death(1). In the thymus, developing T lymphocytes (thymocytes) bearing a T-cell receptor (TCR)-CD3 complex that engages self-antigens are induced to undergo programmed cell death (apoptosis)(2-4), but the mechanisms ensuring this 'negative selection' are unclear. We now report that thymocytes lacking the pro-apoptotic Bcl-2 family member Bim(5,6) (also known as Bcl2l11) are refractory to apoptosis induced by TCR-CD3 stimulation. Moreover, in transgenic mice expressing autoreactive TCRs that provoke widespread deletion, Bim deficiency severely impaired thymocyte killing. TCR ligation upregulated Bim expression and promoted interaction of Bim with Bcl-X-L, inhibiting its survival function. These findings identify Bim as an essential initiator of apoptosis in thymocyte-negative selection.
Publisher
NATURE PUBLISHING GROUP
Keywords
T-CELL-RECEPTOR; INHIBITS MULTIPLE FORMS; NEGATIVE SELECTION; TRANSGENIC MICE; CLONAL DELETION; EXPRESSION; LYMPHOCYTES; ACTIVATION; SURVIVAL; DEATH
Publisher's Version
https://doi.org/10.1038/415922a
Terms of Use/Rights Notice
Refer to copyright notice on published article.


Creation Date: 2002-02-21 12:00:00
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